Discovered catalytic RNA, for which he won the Nobel Prize in 1989
Pioneered biofilm microbiology
Discovered the telomere clock
Studied natural systems and the response of predators to prey
Ethnobotanist: World expert on airborne and allergenic pollen and famous for targeting medicinal plants in the tropical rain forest.
Shared the 2009 Nobel Prize in medicine for the discovery of telomeres.
